Staff at the Galapagos National Park in Ecuador say Lonesome George, a giant tortoise believed to be the last of its subspecies, has died.

加拉帕戈斯国家公园的工作人员称，世界上最后一只同种的巨型陆龟，孤单乔治，已经死亡。

Scientists estimate he was about 100 years old.

Park officials said they would carry out a post-mortem to determine the cause of his death.

With no offspring and no known individuals from his subspecies left, Lonesome George became known as the rarest creature in the world.

For decades, environmentalists unsuccessfully tried to get the Pinta Island tortoise to reproduce with females from a similar subspecies on the Galapagos Islands.

Park officials said the tortoise was found dead in his corral（畜栏） by his keeper of 40 years, Fausto Llerena.

While his exact age was not known, Lonesome George was estimated to be about 100, which made him a young adult as the subspecies can live up to an age of 200.

Galapagos icon¹

Lonesome George was first seen by a Hungarian scientist on the Galapagos island of Pinta in 1972.

Environmentalists had believed his subspecies (Chelonoidis nigra abingdoni) had become extinct.

Lonesome George became part of the Galapagos National Park breeding programme.

After 15 years of living with a female tortoise from the nearby Wolf volcano, Lonesome George did mate, but the eggs were infertile².

He also shared his corral with female tortoises from Espanola island, which are genetically³ closer to him than those from Wolf volcano, but Lonesome George failed to mate with them.

He became a symbol of the Galapagos Islands, which attract some 180,000 visitors a year.